Ladybug Folds Its Hinfwings Like Origami 折り紙

choi

Saito K el al, Investigation of Hindwing Folding in Ladybird Beetles by Artificial Elytron Transplantation and Microcomputed Tomography. Proceedings of the National Academy of Sciences, _: _ (online publication May 17,2017).
www.pnas.org/cgi/doi/10.1073/pnas.1620612114

My comment:
(a) The lead author is teaching assistant Kazuya SAITŌ 斉藤 一哉 助教 of University of Tokyo.
(b) Japanese-English dictionary:
ten-tō-mushi《てんとう虫; 天道虫; 瓢虫》 【てんとうむし】 (n): "ladybug"  (The "mushi" is Japanese pronunciation for 虫.)

(c)
(i) insect wing
https://en.wikipedia.org/wiki/Insect_wing
("are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings")
(ii) elytron
https://en.wikipedia.org/wiki/Elytron
(from Greek for "sheath, cover,"  plural: elytra; is a modified, hardened forewing)
(iii) Coccinellidae
https://en.wikipedia.org/wiki/Coccinellidae
(a family [which is right above 'genus']; section 1 Etymology)

(d) I read many news reports on this, none of which is helpful in visualization of wing folding. In my view, the best source is the figures and video (which can be better) of the paper itself. The link to the paper  is on the top.   

Click "Full Text" on the top horizontal bar.
(i) There is no need to read the text.
(ii) Go to Figure 1 (C) to (G): the right column (sketch) corresponds to left column (a read ladybug).

In the right column, the bluish shield is a (right) elytron. The bigger bigger (right) hindwing will be folded underneath.  Pay no attention to alphabet soup and all sorts of line (heavy or light) -- except the red (hollow) arrow.

In the left column, heed only the red solid arrow.
(iii) Go straight to Figure 3, where (A) shows how a hindwing folds, looking from abdomen of the ladybug to the back. In the rightmost panel, the back (grey in color) folded lengthwise.

In Figure 3(B), looks at the white dotted line only, whose orientation is different from that in 3(A).

Then The end of legend to Figure 3 kindly suggest "(also Movie S3).”  Watch that video.