Who Invented Toothbrush?

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Pagan Kennedy, Who Invented That?  (Toothbrush) New York Times Magazine, Feb 17, 2013.
http://www.nytimes.com/2013/02/1 ... hat-toothbrush.html

Note:
(a) toothbrush
http://en.wikipedia.org/wiki/Toothbrush
(section 1 History)
(b) One of the references (Ref  specifically) of Wiki is

Who Invented the Toothbrush and When Was It Invented?  Science Reference Services, Library of Congress, undated (in the category Everyday Mystery)
http://www.loc.gov/rr/scitech/mysteries/tooth.html
("The bristle toothbrush, similar to the type used today, was not invented until 1498 in China. The bristles were actually the stiff, coarse hairs taken from the back of a hog's neck and attached to handles made of bone or bamboo. Boar bristles were used until 1938, when nylon bristles were introduced by Dupont de Nemours")
(c) Ming Dynasty (1368-1644)